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This feature allows for site admins to lock down what email domain a new user needs to have in order to be given a specific role. On a single site if you attempt to create a new user with a role and email that is not in line with the settings set the user will not be created and a notice informing you that you cant create that specific user with that role will be presented. On a multisite, the user would get generated but not added to the site. You would get a notice that the user could not be added to the site.
cc: @tlovett1